The film centers on paleo-archaeologist Richard Leakey who set out to stop the African ivory poachers whose practices nearly put the noble elephant on a path to extinction.
Oscar winner Eric Roth is writing the screenplay,
David Ellison Jolie, Dana Goldberg and Jon Peters will produce.
The pic also reunites her with cinematographer Roger Deakins after the pair did the biopic Unbroken, which opens on Christmas Day.
In a statement, Jolie says: "I've felt a deep connection to Africa and its culture for much of my life, and was taken with Eric's beautiful script about a man drawn into the violent conflict with elephant poachers who emerged with a deeper understanding of man's footprint and a profound sense of responsibility for the world around him."
The hope is to start production on Africa after Jolie wraps directing By The Sea in which she also stars opposite husband Brad Pitt.
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